Below is an abridged curriculum vitae. Full CV and references are available upon request.


16 Divinity Avenue / Biolabs 2077 / Cambridge, MA 02138

donoughe at


Donoughe, S, Kim, C, and Extavour, CG. (2017). High-throughput live-imaging of embryos in microwell arrays using a modular, inexpensive specimen mounting system. bioRxiv. (PDF) (link to bioRxiv)

de Hoop, M., Donoughe, S., and Boyden, E. (2017). What automation can do for you: The benefits and pitfalls of automating your microscopy research. Science, 355.6323, 421-421. (Webinar; slides available for download at link)

Donoughe, S., and Gliem, S. (2016). High-Throughput, Long-Term Live Imaging: Automated Microscopy of Insect Development. Imaging and Microscopy.

Donoughe, S, and Extavour, CG. (2016). Embryonic development of the cricket Gryllus bimaculatus. Developmental biology, 411(1), 140-156. (PDF) (link to journal)

Donoughe S, Nakamura T, Ewen-Campen B, Green DA, Henderson L, and Extavour CG. (2014) BMP signaling is required for the generation of primordial germ cells in an insect. Proceedings of the National Academy of Science 111: 4133-8. (PDF) (link to journal)

Ewen-Campen B, Donoughe S, Clarke DN, and Extavour CG. (2013) Germ cell specification requires zygotic mechanisms rather than germ plasm in a basally branching insect. Current Biology 23: 835-42. NOTE: Recommended by the Faculty of 1000 (PDF) (link to journal)

Donoughe S and DiNardo S. (2011) Dachsous and Frizzled contribute separately to planar polarity in the Drosophila ventral epidermis. Development 138: 2751-9. NOTE: Recommended by the Faculty of 1000 (PDF) (link to journal)

Donoughe S*, Crall JC*, Merz RA, and Combes SA. (2011) Resilin in dragonfly and damselfly wings and its implications for wing flexibility. Journal of Morphology 272(12): 1409-21. (PDF) (link to journal)


2011-present   Harvard University: Ph.D. candidate in the Department of Organismic and Evolutionary Biology

2004-2008   Swarthmore College: B.A. in Biology

Grants, Awards, Fellowships

2016   International Congress of Entomology Student Award, First Place: awarded to graduate student with the best oral presentation, in this case for a talk on Functional Genomics and Transgenesis

2015   Simmons Award: Research grant awarded to graduate students at Harvard University for whom microscopy is an essential component of their research ($1100)

2014, 2015   Certificate of Distinction in Teaching: This award, sponsored by the Derek Bok Center for Teaching and Learning at Harvard University, acknowledges excellence in one’s work with students and a commitment to teaching

2012   National Science Foundation (NSF) Graduate Research Fellowship ($30,000 x 3 years)

2011-2013   James Mills Pierce Fellowship, Harvard University ($18,000)

2008   Leo M. Leva Prize: Awarded to graduating Biology Majors at Swarthmore College who show unusual promise in the field and whose participation in coursework and departmental activities is noteworthy ($500)

2007   The Norman Meinkoth Field Biology Award: To support summer research on the biomechanics of dragonfly and damselfly wings at Swarthmore College ($1500).

selected Presentations

09/2016   Oral: “Using long-term cell tracking and transgenic crickets to understand how a well-ordered insect embryo is built.” International Congress of Entomology, Orlando, FL. Student competition first place prize.

08/2015   Poster: “Fluid mechanics and collective movement within an egg: Using light sheet microscopy and long term cell tracking to understand how a well-ordered insect embryo is built.” Society for Integrative and Comparative Biology Annual Meeting, Portland, OR.

08/2015   Poster: “High temporal resolution microscopy reveals nuclear dynamics of the insect blastoderm.” Pan-American Society for Evolutionary Developmental Biology Annual Meeting, Berkeley, CA.

03/2015   Oral: “Building a bouncing baby cricket: Early nuclear movements and divisions in Gryllus bimaculatus.” Cellular Dynamics Research Seminar Series. Harvard University, Cambridge, MA. Nominated speaker.

09/2014   Poster: “Understanding the role of BMP signaling in cricket germ cell specification.” Harvard Molecular and Cellular Biology Retreat, Marine Biological Laboratory, Woods Hole, NY.

03/2014   Oral: “Using RNAi to understand BMP signaling and germ cell specification in crickets.” RNAi and Genome Editing Symposium. Tokushima University, Tokushima, Japan. Invited speaker.

10/2012   Poster: “Towards direct evidence for inductive germ cell specification in a basally branching insect, the cricket Gryllus bimaculatus.” Cold Spring Harbor Germ Cell Meeting, Cold Spring Harbor, NY. 

09/2010   Oral: Planar polarity in the Drosophila larval epidermis. Department of Cell and Developmental Biology Chalk Talk Series. University of Pennsylvania School of Medicine, Philadelphia, PA. Nominated speaker.

01/2008   Poster: “Checks on the flex: Structures for maintaining flexible corrugation in odonate wings.” Society for Integrative and Comparative Biology Annual Meeting, San Antonio, TX.

teaching and mentoring

2016   Instructor for “The Beasts Within: How we fight and cooperate with the microscopic organisms living in our bodies,” Harvard University, Cambridge, MA.

Co-leader for a public course on symbiosis and parasitism. Students learn how symbioses evolve, and how they affect our health and the ecosystems around us.

2015–present   Resident Science Tutor, Adams House, Harvard University, Cambridge, MA.

Live alongside Harvard College students, creating and maintaining a safe and healthy living environment, advising students on academic matters, mentoring students that are interested in pursuing careers in the natural sciences. Lead a weekly event called STEM Forum, in which visiting scientists discuss their research with students.

2015   Teaching Fellow for “OEB 57: Animal Behavior,” Harvard University, Cambridge, MA.

Led weekly discussion sections, hosted problem set reviews, and designed and graded assessments. This is an undergraduate-level course offered by the Department of Organismic and Evolutionary Biology. Awarded Certificate of Distinction in Teaching for this course.

2013   Teaching Fellow for “MCB 291: Genetics, Genomics, and Evolution,” Harvard University, Cambridge, MA. 

Designed and graded problem sets, wrote and graded exams, led weekly discussion sections, and led review sessions. This is a graduate-level course offered by the Department of Molecular and Cellular Biology. Awarded Certificate of Distinction in Teaching for this course.

2006-2008   Lab Teaching Assistant, Biology Department, Swarthmore College, Swarthmore, PA

2008   Biology and Organic Chemistry Tutor, Swarthmore College, Swarthmore, PA

2007   Assistant Biology Teacher, Science for Kids, Swarthmore College, Swarthmore, PA